554 554 5.2.3 error when sending SOME emails to my zoho mailbox

554 554 5.2.3 error when sending SOME emails to my zoho mailbox

My client sent screenshot showing that their test email into a Zoho mailbox failed. See attached. Error code: 554 554 5.2.3
However we have tested sending and receiving and encountered no issue, so the account wouldn't seem to be blocked.

So; what could be causing this intermittent error please?