Mails forwarded from GMail often get rejected (541 5.4.1 Mail rejected by destination domain)

Mails forwarded from GMail often get rejected (541 5.4.1 Mail rejected by destination domain)

Hi,

I use Zoho Mail for my domain matczak.org.pl. One of the users set up e-mail forwarding from her GMail account to Zoho account. For a few months now some mails get delivered and some get rejected. There is no apparent key as to what gets delivered and what rejected. Many times e-mails from the same sender (Facebook notifications or an online shop status updates, for example) get delivered once and rejected a couple of minutes/hours later.

I found that people report this being linked to spam filtering. I tried to reset all the spam filtering data in the domain admin panel. I tried disabling the spam filtering for that account altogether. Nothing helps.

The affected account name is 'monika'.

Here's the bounce she gets for the undelivered mails:

---------- Forwarded message ----------
From: Mail Delivery Subsystem < mailer-daemon@googlemail.com>
Date: Thursday, 19 May 2016
Subject: Dolby Laboratories
To: xxxxxxxxxxxxxxxxxx@gmail.com


Delivery to the following recipient failed permanently:

xxxxxx@matczak.org.pl

Technical details of permanent failure:
Google tried to deliver your message, but it was rejected by the server for the recipient domain matczak.org.pl by mx.zohomail.com. [165.254.167.50].

The error that the other server returned was:
541 5.4.1 Mail rejected by destination domain
------------------------

Can I please ask for some support?

Thanks,
Artur Matczak