Randomly getting error 4401

Randomly getting error 4401

On our website, we have a form that is submitted to Zoho CRM via the API (insertRecord method).  It's been running fine for months, we've had no problems.

Since last Thursday or so, we are randomly getting error code 4401 (Unable to populate data, please check if mandatory value is entered correctly.)  It shows up for maybe 10-20 min, and then the error seemingly resolves itself. Without applying any changes or fixes, the script functions are normal and we do not get an error.

Any ideas on what could cause this inconsistency?

Also, just so that I have my bases covered, what are the mandatory fields in regards to the insertRecord method?